A male elephant with an erection pursuing a female at a fast walk or shuffling run. As he Runs-After the estrous female Walks-Away or Runs-Away, often emitting an Estrous-Roar.
References: Poole 1982: 108-109, 174-175; Moss 1983; Poole 1989b; Poole 1996; Kahl & Armstrong, 2000; Poole & Granli 2003; Poole & Granli 2011. (Full reference list)
This behavioral constellation includes the following behaviors: Estrous-Roar, Penis-Erection, Roar, Run-After, Run-Away, Tail-Raising, Walking, Walk-Toward, Walk-Away and occurs in the following context(s): Courtship