Secretion from the swollen temporal glands of a musth male. This secretion, which creates a dark brown streak down the side of the face, mid-way between ear and eye, may be distinguished from Temporin by its congealed appearance and strong odor. Older, higher-ranking musth males secrete more heavily and persistently from enlarged temporal glands. The swollen glands cause their heads to appear even larger and more impressive.
References: Poole 1982: 48, 60, 61, illustration 4.1; Poole, 1987a: 288 illustration; Poole & Granli 2003. (Full reference list)
This behavior occurs in the following context(s): Advertisement & Attraction