Echo, the matriarch of the EB family, our group of primary study, died Sunday 3 May 2009, aged 64 years old. In her memory we below have compiled some images of and vocalizations by this extraordinary and world renown elephant. Echo was the capable leader of her family for at least 36 years.

Echo and her family have been at the core of many discoveries about the nature of elephants as well as the focus of books and films. Some major documentary films by cinematographer Martyn Colbeck have aired around the world, making Echo a household name. This clip from Kenya's NTV 9:00 o'clock evening news on Sunday 10th May shows the high regard in which all Kenyans held Echo.

As an icon for wild elephants Echo has given visitors, viewers and readers enormous pleasure. Echo's legacy will live on - but she will be deeply missed. We feel very priviledged to have spent so much time in her company - following her, with our Amboseli colleagues, for almost half her life.

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All photos ©ElephantVoices

You will enjoy the sounds below much more if you have earphones or are connected to a sound system!